Tuovinen, V., Ekman, S., Thor, G., Vanderpool, D., Spribille, T., Johannesson, H. (2019): Two basidiomycete fungi in the cortex of wolf lichens. Current Biology DOI: 10.1016/j.cub.2018.12.022 Open Access Link
This paper was featured on the cover as well as in The Atlantic


Resl, P., Fernández-Mendoza, F., Mayrhofer, H., Spribille, T. (2018): The evolution of fungal substrate specificity in a widespread group of crustose lichens. Proceedings of the Royal Academy of Sciences B 285: 20180640. Open Access Link

Spribille, T. (2018): Relative symbiont input and the lichen symbiotic outcome. Current Opinion in Plant Biology 44: 57-63.

Simon, A., Goward, T., Di Meglio, J., Dillman, K., Spribille, T., Goffinet, B. (2018) Sticta torii sp. nov., a remarkable lichen of high conservation priority from northwestern North America. Graphis Scripta 30: 105-114. Link


Lücking, R., Moncada, B., McCune, B., Farkas, E., Goffinet, B., Parker, D., Chaves, J.L., Lőkös, L., Nelson, P.R., Spribille, T., Stenroos, S., Wheeler, T., Yanez-Ayabaca, A., Dillman, K., Gockman, O.T., Goward, T., Hollinger, J., Tripp, E.A., Villella, J., Álvaro-Alba, W.R., Arango, C.J., Cáceres, M.E.S., Coca, L.F., Printzen, C., Rodríguez, C., Scharnagl, K., Rozzi, R., Soto-Medina, E., Yakovchenko, L. (2017): Pseudocyphellaria crocata (Ascomycota: Lobariaceae) in the Americas is revealed to be thirteen species, and none of them is P. crocata. Bryologist 120: 441-500.

Sheard, J., Ezhkin, A., Galanina, I., Himelbrant, D., Kuznetsova, E., Shimizu, A., Stepanchikova, I., Thor, G., Tønsberg, T., Yakovchenko, L., Spribille, T. (2017): The lichen genus Rinodina (Physciaceae, Caliciales) in northeastern Asia. Lichenologist 49: 617-672.

Svensson, M., Ekman, S., Klepsland, J.T., Nordin, A., Thor, G., von Hirschheydt, G., Jonsson, F., Knutsson, T., Lif, M., Spribille, T., Westberg, M. (2017): Taxonomic novelties and new records of Fennoscandian crustose lichens. Mycokeys 25: 51-86.


Spribille, T., Tuovinen, V., Resl, P., Vanderpool, D., Wolinski, H., Aime, M.C., Schneider, K., Stabentheiner, E., Toome-Heller, M., Thor, G., Mayrhofer, H., Johannesson, H., McCutcheon, J.P. (2016): Basidiomycete yeasts in the cortex of ascomycete macrolichens. Science 353: 488-492. Link
This paper was featured on the cover as well as in The New York TimesThe Washington PostThe Atlantic, BBC’s Inside ScienceCBCEl PaisDer Standard, Scientific American’s 60-second Science podcast, Scientific American’s Artful Amoeba blog, GizmodoScience NewsSveriges RadioHelsingin SanomatBild der Wissenschaft and other media
Spribille, T. & Mayrhofer, H. (2016): Cyphobasidiaceae, Cyphobasidiales. Index Fungorum 309: 1. Link

Hafellner, J. & Spribille, T. (2016): Tingiopsidium – the correct name for Vestergrenopsis as currently delimited (Koerberiaceae, Peltigerales). Fritschiana (Graz) 83: 47-50.

Schneider, K., Resl, P., Spribille, T. (2016): Escape from the cryptic species trap: lichen evolution on both sides of a cyanobacterial acquisition event. Molecular Ecology 25: 3453-3468. Open Access Download
Subject of a Perspectives piece by Christoph Scheidegger

Resl, P., Mayrhofer, H., Clayden, S.R., Spribille, T., Thor, G., Tønsberg, T., Sheard, J.W. (2016) Molecular, chemical and species delimitation analyses provide new taxonomic insights into two groups of RinodinaLichenologist 48: 469-488. Open Access Download

Holien, H., Palice, Z., Björk, C.R., Goward, T., Spribille, T. (2016): Lecidea coriacea sp. nov., a lichen species from oldgrowth boreal and montane forests in Europe and North America. Herzogia 29 (2) Teil 1: 412-420.

Spribille, T., Hampton-Miller, C.J., Ahti, T., Dillman, K., Fryday, A.M., Thor, G., Tønsberg, T., Schirokauer, D. (2016): Lichens and lichenicolous fungi of Alaska. Monographs in North American Lichenology (in press)

Spribille, T. & Wagner, V. (2016): The starry breck lichen, a dry grassland species on the brink of extinction, gains IUCN status. Bulletin of the Eurasian Dry Grassland Group 31: 27-28. (popular article link to pdf)


Bendiksby, M., Haugan, R., Spribille, T., Timdal, E. (2015): Molecular phylogenetics and taxonomy of the Calvitimela aglaea complex (Tephromelataceae, Lecanorales). Mycologia 107: 1172-1183. Download

Resl, P., Schneider, K., Westberg, M., Printzen, C., Palice, Z., Thor, G., Fryday, A., Mayrhofer, H., Spribille, T. (2015): Diagnostics for a troubled backbone: testing topological hypotheses of trapelioid lichenized fungi in a large-scale phylogeny of Ostropomycetidae (Lecanoromycetidae). Fungal Diversity 73: 239-258. Open Access download

Schneider, K., Resl, P., Westberg, M., Spribille, T. (2015): A new, highly effective primer pair to exclude algae when amplifying nuclear large ribosomal subunit (LSU) DNA from lichens. Lichenologist 47: 269-275.

Spribille, T., Bilovitz, P., Printzen, C., Haugan, R. & Timdal, E. (2015): Buellia asterella. The IUCN Red List of Threatened Species 2015: e.T70385861A70385867. Go to report


Spribille, T., Resl, P., Ahti, T., Pérez-Ortega, S., Mayrhofer, H., Lumbsch, H.T. (2014): Molecular systematics of the wood-inhabiting, lichen-forming genus Xylographa (Baeomycetales, Ostropomycetidae) with eight new species. Symbolae Botanicae Upsalienses 37(1): 1-87. download

McCune, B., Rosentreter, R., Spribille, T., Breuss, O., Wheeler, T. (2014): Montana lichens: an annotated list. Monographs in North American Lichenology 2: 1-183. Order here

Spribille, T., Tønsberg, T., Stabentheiner, E., Muggia, L. (2014): Reassessing evolutionary relationships in the filamentous cyanolichen genus Spilonema (Peltigerales, Lecanoromycetes). Lichenologist 46: 373-388. download

Muggia, L., Pérez-Ortega, S., Fryday, A., Spribille, T., Grube, M. (2014): Global assessment of genetic variation and phenotypic plasticity in the lichen-forming species Tephromela atraFungal Diversity 64: 233-251. download

Wagner, V., Spribille, T., Abrahamczyk, S., Bergmeier, E. (2014) Timberline meadows along a 1000 km transect in NW North America: species diversity and community patterns. Applied Vegetation Science 17: 129-141. download
Featured on the cover


Dickoré, W.B., Schloeder, C., Jacobs, M., Spribille, T., Freitag, H. (2013) Ochotonophila flava (Caryophyllaceae), a new species from Afghanistan. Edinburgh Journal of Botany 70: 405-412. download

Palice, Z., Printzen, C., Spribille, T., Svensson, M., Tønsberg, T., Urbanavichene, I., Yakovchenko, L.S., Ekman, S. (2013) Taxonomy of the genus Myrionora, with a second species from South America. Lichenologist 42: 159-167. download

Spribille, T. & Muggia, L. (2013) Expanded taxon sampling disentangles evolutionary relationships and reveals a new family in Peltigerales (Lecanoromycetes, Ascomycota). Fungal Diversity 58: 171-184. download

Leavitt, S.D., Esslinger, T.L., Spribille, T., Divakar, P.K., Lumbsch, H.T. (2013): Multilocus phylogeny of the lichen-forming fungal genus Melanohalea (Parmeliaceae, Ascomycota): insights on diversity, distributions and a comparison of species tree and concatenated topologies. Molecular Phylogenetics and Evolution 66: 138-152. download


Dillman, K.D., Ahti, T., Björk, C.R., Clerc, P., Ekman, S., Goward, T., Hafellner, J., Pérez-Ortega, S., Printzen, C., Savic, S., Schultz, M., Svensson, M., Thor, G., Tønsberg, T., Vitikainen, O., Westberg, M. & Spribille, T. (2012): New records, range extensions and nomenclatural innovations for lichens and lichenicolous fungi from Alaska, U.S.A.. Herzogia 25: 177-210. download

Sheard, J., Lendemer, J., Spribille, T., Thor, G., Tønsberg, T. (2012) Furthern contributions to the genus Rinodina (Physciaceae, Lecanoromycetidae) in North America: two species new to science and a new record for North America. Herzogia 25: 125-143. download

Grube, M. & Spribille, T. (2012): Exploring symbiont management in lichens. Molecular Ecology 21: 3098-3099. download

Goward, T., Spribille, T., Ahti, T., Hampton-Miller, C. J. (2012): Four new sorediate species in the Hypogymnia austerodes group (lichens) from northwestern North America, with notes on thallus morphology. Bryologist 115:


Muggia, L., Nelson, P., Wheeler, T., Yakovchenko, L.S., Tonsberg, T. & Spribille, T. (2011): Convergent evolution of a symbiotic duet: the case of the lichen genus Polychidium (Peltigerales, Ascomycota). American Journal of Botany 98: 1647-1656. download

Palice, Z., Printzen, C., Spribille, T. & Elix, J.A. (2011): Notes on the synonyms of Lecanora filamentosaGraphis Scripta 23: 1–7. download

Nelson, P.R., Walton, J., Root, H. & Spribille, T. (2011): Hypogymnia pulverata (Parmeliaceae) and Collema leptaleum (Collemataceae), two macrolichens new to Alaska. North American Fungi 6: 1-8. download

Spribille, T., Goffinet, B., Klug, B., Muggia, L., Obermayer, W. & Mayrhofer, H. (2011): Molecular support for the recognition of the Mycoblastus fucatus group as the new genus Violella (Tephromelataceae, Lecanorales). Lichenologist 43: 445-466. download

Spribille, T., B. Klug, H. Mayrhofer (2011): A phylogenetic analysis of the boreal lichen Mycoblastus sanguinarius (Mycoblastaceae, lichenized Ascomycota) reveals cryptic clades correlated with fatty acid profiles. Molecular Phylogenetics and Evolution 59: 603-614. download

Pérez-Ortega, S., Etayo, J. & T. Spribille (2011): A new species of Llimoniella (Ascomycota, Heliotales) on Ramboldia cinnabarina from Alaska. Lichenologist 43: 363–366. download

Galanina, I.A., Yakovchenko, L.S., Tsarenko, N.S., & T. Spribille (2011): Notes on Rinodina excrescens in the Russian Far East (Physciaceae, lichenized Ascomycota). Herzogia 24: 59–64. download

Pérez-Ortega, S., Spribille, T., Palice, Z., Elix, J.A. & C. Printzen (2011): Erratum: A molecular phylogeny of the Lecanora varia group, including a new species from western North America. Mycological Progress 10: 129-130. download

Pérez-Ortega, S. & T. Spribille (2011): Diversidad espectacular de líquenes en un parque nacional de Alaska. Quercus 300: 36-37 (popular article)

Lumbsch, H.T., T. Ahti, S. Altermann, G. Amo De Paz, A. Aptroot, U. Arup, A. Bárcenas Peña, P. A. Bawingan, M. N. Benatti, L. Betancourt, C. R. Björk, K. Boonpragob, M. Brand, F. Bungartz, M. E. S. Cáceres, M. Candan, J. L. Chaves, P. Clerc, R. Common, B. J. Coppins, A. Crespo, M. Dal-Forno, P. K. Divakar, M. V. Duya, J. A. Elix, A. Elvebakk, J. D. Fankhauser, E. Farkas, L. Itatí Ferraro, E. Fischer, D. J. Galloway, E. Gaya, M. Giralt, T. Goward, M. Grube, J. Hafellner, J. E. Hernández M., M. De Los Angeles Herrera Campos, K. Kalb, I. Kärnefelt, G. Kantvilas, D. Killmann, P. Kirika, K. Knudsen, H. Komposch, S. Kondratyuk, J. D. Lawrey, A. Mangold, M. P. Marcelli, B. Mccune, M. I. Messuti, A. Michlig, R. Miranda González, B. Moncada, A. Naikatini, M. P. Nelsen, D. O. Øvstedal, Z. Palice, K. Papong, S. Parnmen, S. Pérez-Ortega, C. Printzen, V. J. Rico, E. Rivas Plata, J. Robayo, D. Rosabal, U. Ruprecht, N. Salazar Allen, L. Sancho, L. Santos De Jesus, T. Santos Vieira, M. Schultz, M. R. D. Seaward, E. Sérusiaux, I. Schmitt, H. J. M. Sipman, M. Sohrabi, U. Søchting, M. Z. Søgaard, L. B. Sparrius, A. Spielmann, T. Spribille, J. Sutjaritturakan, A. Thammathaworn, A.Thell, G. Thor, H. Thüs, E. Timdal, C. Truong, R. Türk, L. Umaña Tenorio, D. K. Upreti, P. van den Boom, M. V. Rebuelta, M. Wedin, S. Will-Wolf, V. Wirth, N. Wirtz, R. Yahr, K. Yeshitela, F. Ziemmeck, T. Wheeler, & R. Lücking (2011): One hundred new species of lichenized fungi: a signature of undiscovered global diversity. Phytotaxa 18: 1–127. download


Crespo, A., Kauff, F., Divakar, P. K., del Prado, R., Pérez-Ortega, S., Amo de Paz, G., Ferencova, Z., Blanco, O., Roca-Valiente, B., Núñez-Zapata, J., Cubas, P., Argüello, A., Elix, J. A., Esslinger, T. L., Hawksworth, D. L., Millanes, A., Molina, C., Wedin, M., Ahti, T., Aptroot, A., Barreno, E., Bungartz, F., Calvelo, S., Candan, M., Cole, M., Ertz, D., Goffinet, B., Lindblom, L., Lücking, R., Lutzoni, F., Mattsson, J.-E., Messuti, M. I., Miadlikowska, J., Piercey-Normore, M., Rico, V. J., Sipman, H. J. M., Schmitt, I., Spribille, T., Thell, A., Thor, G., Upreti, D. K., & Lumbsch, H. T. (2010): Phylogenetic generic classification of parmelioid lichens (Parmeliaceae, Ascomycota) based on molecular, morphological and chemical evidence. Taxon 59: 1735-1753. download
Spribille, T., Pérez-Ortega, S., Tønsberg, T., Schirokauer, D. (2010): Lichens and lichenicolous fungi of the Klondike Gold Rush National Historic Park, Alaska, in a global biodiversity context. Bryologist 113: 439-515. download
This paper was featured by Science DailyThe Juneau Empire, and Alaska Public Radio Network as well as on the cover.

Schmitt, I., Fankhauser, J.D., Sweeney, K., Spribille, T., Kalb, K., Lumbsch, H.T. (2010): Gyalectoid Pertusaria species form a sister-clade to Coccotrema (Ostropomycetidae, Ascomycota) and comprise the new lichen genus Gyalectaria. Mycology 1: 75-83. download

Pérez Ortega, S., Spribille, T., Palice, Z., Elix, J.A. & Printzen, C. (2010): A molecular phylogeny of the Lecanora varia group, including a new species from western North America. Mycological Progress 9: 523-535. download

Spribille, T. & H.T. Lumbsch (2010): A new species of Elixia (Umbilicariales) from Greece. Lichenologist 42: 365-371.  download

Goward, T., T. Ahti, J.A. Elix & T. Spribille (2010): Hypogymnia recurva and Hypogymnia wilfiana spp. nov., two new lichens from western North America. – Botany/Botanique 88: 345-351. download


Pérez-Ortega, S. & T. Spribille (2009): Lepraria torii, a new epiphytic species with fumarprotocetraric acid from northwest North America. Graphis Scripta 21: 36-41. download

Björk, C.R., Goward, T., Spribille, T. (2009): New records and range extensions of rare lichens from waterfalls and sprayzones in inland British Columbia, Canada. Evansia 26: 219-224. download

Etayo, J., Palice, Z., Spribille, T. (2009): Candelariella boleana, a new epiphytic species from southern and central Europe (Candelariaceae, Ascomycota). Nova Hedwigia 89(3-4): 545-552. download

Pérez-Ortega, S., Spribille, T. (2009): Three new species of Lichenopeltella (Microthyriaceae, Ascomycota) from northwest North America. Nova Hedwigia 89 (1-2): 219-228.  download

Spribille, T. (2009): Fuscopannaria leucosticta (Tuck.) P.M. Jørg. rediscovered in Europe. Lichenologist 41: 209–210. download

Heininger, C. & Spribille, T. (2009): The sorediate species of Xylographa in Austria (Baeomycetales, lichenized Ascomycetes). Herzogia 22: 129-134. download

Spribille, T., Björk, C.R., Ekman, S., Elix, J.A., Goward, T., Printzen, C., Tønsberg, T., Wheeler, T. (2009): Contributions to an epiphytic lichen flora of northwest North America. I. Eight new species from British Columbia inland rainforests. Bryologist 112: 109–137. download
Pre-featured on the March 29, 2007 cover of the Vancouver Sun.


Spribille, T., Björk, C.R. (2008): New records and range extensions in the North American lignicolous lichen flora. Mycotaxon 105: 455–468. download

Spribille, T., Thor, G., Bunnell, F.L., Goward, T., Björk, C.R. (2008): Lichens on dead wood: species-substrate relationships in the epiphytic lichens floras of the Pacific Northwest and Fennoscandia. Ecography 31: 741-750. download

Printzen, C., Spribille, T., Tønsberg, T. (2008): Myochroidea, a new genus of corticolous, crustose lichens to accomodate the Lecidea leprosula group. Lichenologist 40: 195–207.  download

Schmull, M., Spribille, T. 2007. Schaereria. pp. 396-397 in T. H. Nash III, C. Gries, F. Bungartz, eds. Lichen Flora of The Greater Sonoran Region, Vol. III. Lichens Unlimited, Tempe, Arizona.

Bunnell, F.L., Spribille, T., Houde, I., Goward, T., Björk, C. (2008): Lichens on down wood in logged and unlogged forest stands. Canadian Journal of Forest Research 38: 1033-1041. download


Houde, I., Leech, S., Bunnell, F.L., Spribille, T. & Bjork, C. (2007): Old forest remnants contribute to sustaining biodiversity: the case of the Albert River valley. British Columbia Journal of Ecosystems and Management 8 (3): 43-52. download

Pérez-Ortega, S., Spribille, T.(2007): Bellemerella ritae sp. nov. (Verrucariaceae), a new lichenicolous ascomycete from northwest North America.Nova Hedwigia 85(3-4): 512-520. download

Spribille, T., Jørgensen, P.M., Schultz, M., Houde, I. (2007): Santessoniella saximontana, a new lichen species from the Rocky Mountains. Bibliotheca Lichenologica 96: 287-297. download

Spribille, T. & C. Printzen (2007): Lecidea rubrocastanea, a new lichen species from conifer bark and wood in interior western North America (Lecanorales, lichenized Ascomycetes). Lichenologist 39: 339-347. download

Spribille, T. & T. Tønsberg (2007): Lepraria bergensis and L. obtusatica new to Germany. Herzogia 20: 327-328. download

Thiel, H. & T. Spribille (2007): Lichens and bryophytes on shaded sandstone outcrops used for rock climbing in the vicinity of Göttingen (southern Lower Saxony, Germany). Herzogia 20: 159-177. download

Spribille, T. & T. Tønsberg (2007): Mycoblastus marginatus, a new synonym for M. affinis (Mycoblastaceae, lichenized Ascomycota). Mycotaxon 100: 105-107.


Hauck, M., Paul, A. & Spribille, T. (2006): Uptake and toxicity of manganese in epiphytic cyanolichens. Environmental and Experimental Botany 53: 216-224. download

Spribille, T., Schultz, M., Breuss, O., Bergmeier, E. (2006): Notes on the lichens and lichenicolous fungi of western Crete (Greece). Herzogia 19: 125-148. download


Goward, T. & Spribille, T. (2005): Lichenological evidence for the recognition of inland rainforests in western North America. Journal of Biogeography 32: 1209-1219. download

Hauck, M. & Spribille, M. (2005): The significance of precipitation and substrate chemistry for epiphytic lichen diversity in spruce-fir forests of the Salish Mountains, northwestern Montana. Flora 200: 547-562. download

Schmull, M. & Spribille, T. (2005): Schaereria dolodes (Nyl. Ex Hasse) Schmull & T. Spribille: a second corticolous species in the genus.  Lichenologist 37: 527-533. download


Coppins, B.J. & Spribille, T. (2004): Micarea subalpina Coppins & Spribille, a new subalpine species from the Rocky Mountains. Lichenologist 36: 97-102.


Spribille, T. & Hauck, M. (2003): Pyrrhospora gowardiana, a new montane lichen from western North America (Lecanoraceae, lichenized ascomycetes). Bryologis106: 560-564.


Hauck, M. & Spribille, T. (2002): The Mn/Ca and Mn/Mg ratios in bark as possible causes for the occurrence of Lobarion lichens on conifers in the dripzone of Populus in western North America. Lichenologist 34: 527-532. download

Spribille, T. (2002): Additions to the lichen flora of Alberta, Canada from Crowsnest Pass. Evansia 19: 20-21.

Spribille, T. (2002): Pilophorus clavatus and Pseudocyphellaria crocata new to the interior of northwestern North America. Evansia 19: 81-82. download

Spribille, T.; Heidel, B.; Albert, W.E.; Triepke, F.J.; Vanderhorst, J.; Arvidson, G.M. (2002): Noteworthy collections: Montana. Madroño 49(1): 55-58.

Spribille, T. (2002): Noteworthy collections: Idaho. Madroño 49(1): 54-55.

Spribille, T. (2002): The mountain forests of British Columbia and the American Northwest: floristic patterns and syntaxonomy.  Folia Geobotanica 37: 475-508. –download

Spribille, T. & Chytrý, M. (2002): Floristic vegetation surveys in the circumboreal coniferous forests: a review.  Folia Geobotanica 37: 365-382. download

Stachurska-Swakoń, A. & Spribille, T. (2002): Forest communities of the northern Whitefish Range, Rocky Mountains, Montana, U.S.A.  Folia Geobotanica 37: 509-540. download


Breuss, O. & Spribille, T. (2001): Verrucaria kootenaica, a new pyrenocarpous lichen from the northern Rocky Mountains.  Bryologist 104: 453-455.

Kolb, A. & Spribille, T. (2001): Calicium corynellum (Ach.) Ach. in the United States and Calicium montanum Tibell new to North America.  Evansia 18: 90-92. download

Spribille, T., Stroh, H.G. & Triepke, F.J. (2001): Are habitat types compatible with floristically-defined associations?  Journal of Vegetation Science 12: 791-796. download


Elliott, J.C. & Spribille, T. (2000): Montana collections of Scorpidium scorpioides and Meesia triquetra Evansia 17: 10-13.

Spribille, T. & Kolb, A. (2000): Cetraria sepincola new to Idaho and Melanelia septentrionalis new to Montana, with notes on their distribution and ecology.  Evansia 17: 112-115.


Greven, H.E. & Spribille, T. (1999): Grimmia brittoniae, a rare moss endemic to northwestern Montana.  Bryologist 102: 116-118.

Spribille, T. (1999): Forest vegetation in the northern Salish Mountains of northwestern Montana, U.S.A.  Phytocoenologia 29: 503-577.


Spribille, T. (1998): Bryophytes of the Yellow Bay Biological Station, Lake County, Montana.  Evansia 15: 1-4.